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There were giants in the earth in those days; and also after that, when the sons of God came in unto the daughters of men, and they bore children to them. --Genesis 6:5

Sister Evangeline was just a girl when her father entrusted her to the Franciscan Sisters of Perpetual Adoration in upstate New York. Now, at twenty-three, her discovery of a 1943 letter from the famous philanthropist Abigail Rockefeller to the late mother superior of Saint Rose Convent plunges Evangeline into a secret history that stretches back a thousand years: an ancient conflict between the Society of Angelologists and the monstrously beautiful descendants of angels and humans, the Nephilim.

For the secrets these letters guard are desperately coveted by the once-powerful Nephilim, who aim to perpetuate war, subvert the good in humanity, and dominate mankind. Generations of angelologists have devoted their lives to stopping them, and their shared mission, which Evangeline has long been destined to join, reaches from her bucolic abbey on the Hudson to the apex of insular wealth in New York, to the Montparnasse cemetery in Paris and the mountains of Bulgaria.

A decade has passed since Verlaine saw Evangeline alight from the Brooklyn Bridge, the sight of her new wings a betrayal that haunts him still. Now an elite angel hunter for the Society of Angelology, he pursues his mission with single-minded devotion: to capture, imprison, and eliminate her kind.

But when Evangeline suddenly appears on a twilit Paris street, Verlaine finds her nature to be unlike any of the other creatures he so mercilessly pursues, casting him into a spiral of doubt and confusion that only grows when she is abducted before his eyes by a creature who has topped the society's most-wanted list for more than a century. The ensuing chase drives Verlaine and his fellow angelologists from the shadows of the Eiffel Tower to the palaces of St. Petersburg and deep into the provinces of Siberia and the Black Sea coast, where the truth of Evangeline's origins--as well as forces that could restore or annihilate them all--lie in wait.